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Pheidole absurda Forel 1886
Pheidole ajax Forel 1899
Pheidole alfaroi Emery 1896
Pheidole amabilis Wilson 2003
Pheidole anastasii Emery 1896
Pheidole angulifera Wilson 2003
Pheidole arachnion Wilson 2003
Pheidole arboricola Wilson 2003
Pheidole arietans Wilson 2003
Pheidole arhuaca Foreal 1901
Pheidole beloceps Wilson 2003
Pheidole biconstricta Mayr 1870
Pheidole bicornis Forel 1899
Pheidole bilimeki Mayr 1870
Pheidole biolleyi Forel 1908
Pheidole boliviana Wilson 2003
Pheidole boltoni Wilson 2003
Pheidole boruca Wilson 2003
Pheidole brachyops Wilson 2003
Pheidole browni Wilson 2003
Pheidole caltrop Wilson 2003
Pheidole carapuna Mann 1916
Pheidole carinata Wilson 2003
Pheidole cataphracta Wilson 2003
Pheidole caulicola Wilson 2003
Pheidole celaena Wilson 2003
Pheidole cephalica F. Smith 1858
Pheidole cerina Wilson 2003
Pheidole chocoensis Wilson 2003
Pheidole christopherseni Forel 1912
Pheidole citrina Wilson 2003
Pheidole cocciphaga Borgmeier 1934
Pheidole colobopsis Mann 1916
Pheidole cramptoni Wheeler 1916
Pheidole daphne Wilson 2003
Pheidole dasypyx Wilson 2003
Pheidole diabolus Wilson 2003
Pheidole diana Forel 1908
Pheidole dossena Wilson 2003
Pheidole dryas Wilson 2003
Pheidole ectatommoides Wilson 2003
Pheidole erratilis Wilson 2003
Pheidole exarata Emery 1896
Pheidole excubitor Wilson 2003
Pheidole fallax Mayr 1870
Pheidole fariasana Wilson 2003
Pheidole fimbriata Roger 1863
Pheidole fiorii Emery 1890
Pheidole flavens Roger 1863
Pheidole gauthieri Forel 1901
Pheidole glomericeps Wilson 2003
Pheidole gouldi Forel 1886
Pheidole gradifer Wilson 2003
Pheidole hasticeps Wilson 2003
Pheidole hazenae Wilson 2003
Pheidole hector Wilson 2003
Pheidole hirsuta Emery 1896
Pheidole hizemops Wilson 2003
Pheidole hoelldobleri Wilson 2003
Pheidole indagatrix Wilson 2003
Pheidole indistincta Forel 1899
Pheidole innupta Menozzi 1931
Pheidole JTL-006 Longino ms
Pheidole JTL-078 Longino ms
Pheidole JTL-104 Longino ms
Pheidole JTL-110 Longino ms
Pheidole JTL-112 Longino ms
Pheidole JTL-113 Longino ms
Pheidole JTL-122 Longino and Cover ms
Pheidole JTL-123 Longino and Cover ms
Pheidole JTL-125 Longino and Cover ms
Pheidole JTL-126 Longino and Cover ms
Pheidole JTL-129 Longino and Cover ms
Pheidole JTL-130 Longino and Cover ms
Pheidole JTL-132 Longino ms
Pheidole JTL-133 Longino ms
Pheidole JTL-134 Longino ms
Pheidole JTL-135 Longino ms
Pheidole JTL-136 Longino ms
Pheidole JTL-137 Longino ms
Pheidole JTL-138 Longino ms
Pheidole JTL-139 Longino ms
Pheidole JTL-140 Longino ms
Pheidole JTL-141 Longino ms
Pheidole JTL-142 Longino ms
Pheidole JTL-143 Longino ms
Pheidole JTL-144 Longino ms
Pheidole JTL-153 Longino ms
Pheidole JTL-154 Longino ms
Pheidole JTL-157 Longino ms
Pheidole kukrana Wilson 2003
Pheidole laselva Wilson 2003
Pheidole laticornis Wilson 2003
Pheidole longinoi Wilson 2003
Pheidole lourothi Wilson 2003
Pheidole lucaris Wilson 2003
Pheidole lustrata Wilson 2003
Pheidole mantilla Wilson 2003
Pheidole megacephala (Fabricius 1793)
Pheidole mendicula Wheeler 1925
Pheidole monteverdensis Wilson 2003
Pheidole multispina Wilson 2003
Pheidole nasutoides Holldobler and Wilson 1992
Pheidole nebulosa Wilson 2003
Pheidole nigricula Wilson 2003
Pheidole nitella Wilson 2003
Pheidole nitidicollis Emery 1896
Pheidole olsoni Wilson 2003
Pheidole onyx Wilson 2003
Pheidole otisi Wilson 2003
Pheidole perpusilla Emery 1894
Pheidole potosiana Wilson 2003
Pheidole prattorum Wilson 2003
Pheidole prolixa Wilson 2003
Pheidole prostrata Wilson 2003
Pheidole protensa Wilson 2003
Pheidole psilogaster Wilson 2003
Pheidole pugnax Dalla Torre 1892
Pheidole punctatissima Mayr 1870
Pheidole radoszkowskii Mayr 1884
Pheidole rectispina Wilson 2003
Pheidole rectitrudis Wilson 2003
Pheidole renae Wilson 2003
Pheidole rhinoceros Forel 1899
Pheidole rogeri Emery 1896
Pheidole rugiceps Wilson 2003
Pheidole ruida Wilson 2003
Pheidole sagittaria Wilson 2003
Pheidole scalaris Wilson 2003
Pheidole scitula Wilson 2003
Pheidole scrobifera Emery 1896
Pheidole sensitiva Borgmeier 1959
Pheidole sicaria Wilson 2003
Pheidole simonsi Wilson 2003
Pheidole spathipilosa Wilson 2003
Pheidole specularis Wilson 2003
Pheidole striaticeps Mayr 1870
Pheidole stulta Forel 1886
Pheidole subarmata Mayr 1884
Pheidole susannae Forel 1886
Pheidole synarmata Wilson 2003
Pheidole tanyscapa Wilson 2003
Pheidole tennantae Wilson 2003
Pheidole traini Wilson 2003
Pheidole truncula Wilson 2003
Pheidole ulothrix Wilson 2003
Pheidole umphreyi Wilson 2003
Pheidole ursus Mayr 1870
Pheidole vallifica Forel 1901
Pheidole variegata Emery 1896
Pheidole veletis Wilson 2003
Pheidole verricula Wilson 2003
Pheidole vestita Wilson 2003
Pheidole violacea Wilson 2003
Many of the names on the Pheidole of Costa Rica webpages changed with the publication of Wilson (2003). Below is a summary of those changes.
Old name = New name
campanae = tennantae
decem = perpusilla
JTL-009 = brachyops
JTL-013 = hoelldobleri
JTL-021 = truncula
JTL-036 = glomericeps
JTL-040 = amabilis
JTL-044 = ulothrix
JTL-046 = dryas
JTL-048 = erratilis
JTL-050 = carinata
JTL-051 = erratilis
JTL-052 = cataphracta
JTL-054 = caulicola
JTL-066 = hector
JTL-068 = chocoensis
JTL-073 = mantilla
JTL-074 = violacea
JTL-077 = psilogaster
JTL-079 = laticornis
JTL-080 = potosiana
JTL-081 = vestita
JTL-083 = cerina
JTL-087 = renae
JTL-090 = lustrata
JTL-091 = arboricola
JTL-092 = lucaris
JTL-094 = daphne
JTL-095 = hizemops
JTL-097 = longinoi
JTL-099 = onyx
JTL-100 = boliviana
JTL-102 = beloceps
JTL-103 = prolixa
JTL-106 = arhuaca
JTL-111 = colobopsis
JTL-114 = scitula
JTL-115 = hizemops
JTL-116 = angulifera
JTL-118 = excubitor
JTL-119 = exarata
JTL-121 = perpusilla
JTL-127 = synarmata
JTL-128 = hazenae
JTL-145 = synarmata
pubiventris = variegata
praetorum = prattorum
radoszkowskii luteola = radoszkowskii
radoszkowskii pugnax = pugnax
sus = traini
There was early confusion in the names of two species: rectitrudis vs. erectitrudis and rectispina vs. erectispina. In Wilson (2003) they are rectitrudis and rectispina.
Wilson aptly describes Pheidole as a "hyperdiverse" ant genus. Local communities in the Neotropics can contain over 70 species, and there is extensive geographic variation. There will be on-going assessment of whether this variation is inter- versus intra-specific, and thus Pheidole species names will undoubtedly remain unstable for a while.
The history of Pheidole taxonomy for Costa Rica begins with the early naturalists. Alfaro, Tristan, Biolley, and others collected in Costa Rica in the late 1800's and sent material to European taxonomists, mainly Emery and Forel. From about 1880 to 1920 Emery and Forel described many new species from this material, making Costa Rica the type locality for many Neotropical Pheidole species. Starting in the 1960's, Bill Brown, E. O. Wilson, and others began collecting ants in Costa Rica, accumulating material in collections at Harvard's Museum of Comparative Zoology (MCZ). Costa Rica subsequently became a mecca for students of tropical biology, attracting a large number of researchers, many of whom studied ants. During this time I became interested in the ant fauna of Costa Rica and began collecting and sorting Pheidole material. Meanwhile E. O. Wilson and Stefan Cover of the MCZ began an ambitious project to revise the Pheidole of the New World. In 1997 Stefan Cover and I collaborated to develop the Pheidole pages for the Ants of Costa Rica website. These pages used a combination of older published names, Wilson's unpublished manuscript names, and a series of morphospecies codes of the form "JTL-000." Wilson's revision was published in 2003 (Wilson, E. O. 2003. Pheidole in the New World: A Dominant, Hyperdiverse Ant Genus. Harvard University Press, Cambridge, Mass).
There are inevitable differences of opinion in taxonomy, and below I attempt to outline some of the differences between the Pheidole taxonomy presented here and the taxonomy in Wilson's revision.
The following are potential synonyms (Junior syn. = senior syn.):
angusticeps = gradifer
iracunda = arachnion
chloe = striaticeps
ebenina = laselva
exigua = flavens
glyphoderma = arietans
grantae = exarata
humida = boltoni
indagatrix = obscurior
medialis = radoszkowskii
perdiligens = indistincta
petersoni = erratilis
piceonigra = indistincta
sagana = nitidicollis
texticeps = indistincta
Below are Wilson synonyms I disagree with:
innupta (synonymized under alfaroi)
anastasii (synonymized under bilimeki)
Page author: John T. Longino longinoj@evergreen.edu
Date of this version: 29 December 2008.
Previous versions of this page: 27 December 1997, 25 November 1998, 29 January 2001, 29 June 2003
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