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Pheidole species names will undoubtedly remain unstable for a while. Many Neotropical species were described from about 1880 to 1920, by Emery and Forel. Alfaro, Tristan, Biolley, and others collected in Costa Rica and sent material to European taxonomists, and thus Costa Rica is the type locality for many Neotropical species. The types reside in European museums and thus there is an inevitable delay in comparing newly delimited species with types.
The species names in the list exhibit varying degrees of reliability. The most reliable names are usually the published names whose type locality is Costa Rica or the adjacent parts of Panama. Published names whose type localities are more distant may be less reliable. E. O. Wilson is carrying out major revisionary work on the genus, and the early parts of this work were done in collaboration with W. L. Brown, Jr. Thus there are many unpublished manuscript names of Wilson and Brown. These are mostly undescribed species, but a few will probably turn out to have prior names. The species given a code number of the form "JTL-000" are newly delimited species whose status as new versus previously described is unknown (see discussion of names).
I have not examined types of the following taxa, all of which have type localities in Costa Rica:
biconstricta surda Forel 1912:222
biconstricta bicolor Emery 1890:50
biconstricta bicolor regina (unavailable)
biconstricta rubicunda Emery 1890:50
biolleyi tristani Forel 1908:50
cephalica incrustata Forel 1908:59
cephalica sarrita Forel 1908:59
crassipes grantae Forel 1908:53
dimidiata nitidicollis Emery 1896:79
exarata Emery 1896:68
plebecula Forel 1899:68
pubiventris variegata Emery 1896:72
Pheidole absurda Forel 1886
Pheidole ajax Forel 1899
Pheidole alfaroi Emery 1896
Pheidole anastasii Emery 1896
Pheidole arachnion Wilson 2003
Pheidole arietans Wilson 2003
Pheidole biconstricta Mayr 1870
Pheidole bicornis Forel 1899
Pheidole biolleyi Forel 1908
Pheidole boruca Wilson 2003
Pheidole caltrop Wilson 2003
Pheidole campanae Wilson 2003
Pheidole carapuna Mann 1916
Pheidole celaena Wilson 2003
Pheidole cephalica F. Smith 1858
Pheidole christopherseni Forel 1912
Pheidole citrina Wilson 2003
Pheidole cocciphaga Borgmeier 1934
Pheidole cramptoni Wheeler 1916
Pheidole dasypyx Wilson 2003
Pheidole decem Forel 1901
Pheidole diabolus Wilson 2003
Pheidole diana Forel 1908
Pheidole dossena Wilson 2003
Pheidole ectatommoides Wilson 2003
Pheidole fallax Mayr 1870
Pheidole fimbriata Roger 1863
Pheidole fiorii Emery 1890
Pheidole flavens Roger 1863
Pheidole gauthieri Forel 1901
Pheidole gouldi Forel 1886
Pheidole hirsuta Emery 1896
Pheidole indagatrix Wilson 2003
Pheidole indistincta Forel 1899
Pheidole innupta Menozzi 1931
Pheidole JTL-005 Longino ms
Pheidole JTL-006 Longino ms
Pheidole JTL-009 Longino ms
Pheidole JTL-013 Longino ms
Pheidole JTL-021 Longino ms
Pheidole JTL-027 Longino ms
Pheidole JTL-036 Longino ms
Pheidole JTL-040 Longino ms
Pheidole JTL-044 Longino ms
Pheidole JTL-046 Longino ms
Pheidole JTL-048 Longino ms
Pheidole JTL-050 Longino ms
Pheidole JTL-051 Longino ms
Pheidole JTL-052 Longino ms
Pheidole JTL-054 Longino ms
Pheidole JTL-066 Longino ms
Pheidole JTL-068 Longino ms
Pheidole JTL-070 Longino ms
Pheidole JTL-073 Longino ms
Pheidole JTL-074 Longino ms
Pheidole JTL-077 Longino ms
Pheidole JTL-078 Longino ms
Pheidole JTL-079 Longino ms
Pheidole JTL-080 Longino ms
Pheidole JTL-081 Longino ms
Pheidole JTL-087 Longino ms
Pheidole JTL-090 Longino ms
Pheidole JTL-091 Longino ms
Pheidole JTL-092 Longino ms
Pheidole JTL-094 Longino ms
Pheidole JTL-095 Longino ms
Pheidole JTL-097 Longino ms
Pheidole JTL-099 Longino ms
Pheidole JTL-100 Longino ms
Pheidole JTL-102 Longino ms
Pheidole JTL-103 Longino ms
Pheidole JTL-104 Longino ms
Pheidole JTL-106 Longino ms
Pheidole JTL-111 Longino ms
Pheidole JTL-112 Longino ms
Pheidole JTL-113 Longino ms
Pheidole JTL-116 Longino ms
Pheidole JTL-118 Longino and Cover ms (cf. championi)
Pheidole JTL-119 Longino and Cover ms (cf. championi)
Pheidole JTL-121 Longino and Cover ms (cf. decem)
Pheidole kukrana Wilson 2003
Pheidole laselva Wilson 2003
Pheidole megacephala (Fabricius 1793)
Pheidole mendicula Wheeler 1925
Pheidole monteverdensis Wilson 2003
Pheidole multispina Wilson 2003
Pheidole nasutoides Holldobler and Wilson 1992
Pheidole nebulosa Wilson 2003
Pheidole nigricula Wilson 2003
Pheidole nitella Wilson 2003
Pheidole obliquicantha Wilson 2003
Pheidole olsoni Wilson 2003
Pheidole otisi Wilson 2003
Pheidole praetorum Wilson 2003
Pheidole prostrata Wilson 2003
Pheidole psilogaster Wilson 2003
Pheidole pubiventris Mayr 1887
Pheidole punctatissima Mayr 1870
Pheidole punctatissima annectens Wheeler 1905
Pheidole radoszkowskii luteola Forel 1893
Pheidole radoszkowskii pugnax Dalla Torre 1892
Pheidole rectispina Wilson 2003
Pheidole rectitrudis Wilson 2003
Pheidole rhinoceros Forel 1899
Pheidole rogeri Emery 1896
Pheidole rugiceps Wilson 2003
Pheidole ruida Wilson 2003
Pheidole sagana Wheeler 1934
Pheidole sagittaria Wilson 2003
Pheidole scalaris Wilson 2003
Pheidole scrobifera Emery 1896
Pheidole sensitiva Borgmeier 1959
Pheidole sicaria Wilson 2003
Pheidole simonsi Wilson 2003
Pheidole spathipilosa Wilson 2003
Pheidole specularis Wilson 2003
Pheidole striaticeps Mayr 1870
Pheidole subarmata Mayr 1884
Pheidole sus Wilson 2003
Pheidole susannae Forel 1886
Pheidole tanyscapa Wilson 2003
Pheidole umphreyi Wilson 2003
Pheidole verricula Wilson 2003
Pheidole vestita Wilson 2003
Page author: John T. Longino longinoj@evergreen.edu
Date of this version: 27 December 1997
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