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Submitted Manuscripts

Brown, B. V. Preliminary analysis of a host shift: revision of the Neotropical species of Apocephalus, subgenus Mesophora (Diptera: Phoridae). Submitted to Contributions in Science.

In Press

Brehm, G., R. K. Colwell, and J. Kluge. In press. The role of environment and mid-domain effect on moth species richness along a tropical elevational gradient. Global Ecology & Biogeography.

Gillespie, J. J., Cannone, J. J., Gutell, R. R., and Cognato, A. I. in press. A secondary structural model of the 28S rRNA expansion segments D2 and D3 from rootworms and related leaf beetles (Coleoptera: Chrysomelidae; Galerucinae). Insect Molecular Biology.

Karg, W. in press. Die euedaphische Raubmilbengattung Multidentorhodacarus Shcherbak, 1980 mit einer neuen Art aus Mittelamerika [The euedaphic predatory mite genus Multidentorhodacarus Shcherbak, 1980 with a new species from Central America]. Abh. Ber. Naturkundemus. Gorlitz, 72. [M. squamosus in lit.]

Karg, W. in press. Zur Systematik der Raubmilbenfamilien Hypoaspididae v. Vitzthum, 1941 und Rhodacaridae Oudemans, 1902 (Acarina, Parasitiformes) mit neuen Arten aus Sud- und Mittelamerika [To the systematics of the predatory mite families Hypoaspididae v. Vitzthum, 1941 and Rhodacaridae Oudemans, 1902 (Acarina, Parasitiformes) with new species from South- and Central America]. Mitt. Mus. Nat.kd. Berlin, Zool. Reihe, 76. [Oloopticus nudus in lit., O. retiventer in lit., Interrhodeus n. gen. in lit., I. brevicornis in lit.]

Kirkendall, L. R. in press. A new, host-specific Neotropical ambrosia beetle (Curculionidae, Scolytinae, Xyleborus) breeding in live trees. Annals of the Entomological Society of America.

Longino, J. T. In press. Complex nesting behavior by two neotropical species of the ant genus Stenamma (Hymenoptera: Formicidae). Biotropica.

Longino, J. T., and F. Fernández. In press. A taxonomic review of the genus Wasmannia (Hymenoptera: Formicidae: Myrmicinae). Memoirs of the American Entomological Institute.

Nickle, D. A. and P. Naskrecki, In press. A review of the South American genus Loboscelis Karny (Tettigoniidae: Conocephalinae s.l.). Journal of Orthoptera Research. 10 ms. pages.

Smith, D. R. In press. Checklist of the Pergidae (Hymenoptera: Symphyta) of the Western Hemisphere, with a new genus and two new species. In Blank S. M. and A. Taeger, eds. Recent Sawfly Research: Synthesis and Prospects.

Westenberger, S. J., N. R. Sturm, D. Yanega, S. A. Podlipaev, R. Zeledon, D. A. Campbell and D. A. Maslov. in press. Trypanosomatid biodiversity in Costa Rica: genotyping of parasites from Heteroptera using the Spliced Leader RNA gene. Parasitology.


Cardelús, C., R. K. Colwell, and J. E. Watkins, Jr. 2006. Vascular epiphyte distribution patterns: explaining the mid-elevation richness peak. J. Ecology 94:144-156.

Chao, A., R. L. Chazdon, R. K. Colwell, and T.-J. Shen. 2006. Abundance-based similarity indices and their estimation when there are unseen species in samples. Biometrics 62:361-371.

Dunn, R. R., R. K. Colwell, and C. Nilsson. 2006. The river domain: why are there more species halfway up the river? Ecography 29:251-259.

Watkins, J. E., Jr., C. Cardelús, R. C. Moran, and R. K. Colwell. 2006. Diversity and distribution of ferns along an elevational gradient in Costa Rica. American Journal of Botany 93:73-83.


Bunker, D. E., F. DeClerck, J. C. Bradford, R. K. Colwell, I. Perfecto, O. L. Phillips, M. Sankaran, and S. Naeem. 2005 (20 October). Species loss and aboveground carbon storage in a tropical forest. Science (Express Reports) 10.1126/science.1117682:1-5.

Chao, A., R. L. Chazdon, R. K. Colwell, and T.-J. Shen. 2005. A new statistical approach for assessing compositional similarity based on incidence and abundance data. Ecology Letters 8:148-159.

Colwell, R. K., C. Rahbek, and N. Gotelli. 2005. The mid-domain effect: there's a baby in the bathwater. American Naturalist 166:E149-E154.

Davis, D. R. and D. L. Wagner. 2005. Biology and systematics of the neotropical leaf-mining genus Eucosmophora (Lepidoptera: Gracillariidae). Tropical Lepidoptera 13:1-40.

Epstein, M. E., and J. F.Corrales. 2005. Twenty-five new species of Costa Rican Limacodidae (Lepidoptera: Zygaenoidea). Zootaxa 701:1-86.

Hamilton, R. W. 2005. Omolabus Jekel in north and central America (Coleoptera: Attelabidae). Zootaxa 986:1-60.

Hespenheide, H.A. 2005. Weevils of the genera Archocopturus Heller and Zygopsella Champion, sibling species, and mimetic homoplasy (Coleoptera: Curculionidae: Conoderinae). Proceedings of the Entomological Society of Washington. 107: 671-685.

Mao, C. X., R. K. Colwell, and J. Chang. 2005. Estimating species accumulation curves using mixtures. Biometrics 61:433–441.

Mao, C. X., and R. K. Colwell. 2005. Estimation of species richness: mixture models, the role of rare species, and inferential challenges. Ecology Ecology 86:1143-1153.

Opitz, W. 2005. Classification, natural history, and evolution of the genus Aphelocerus Kirsch (Coleoptera: Cleridae: Clerinae). Bulletin of the American Museum of Natural History 293:1-128.

Robbins, R. K., and M. Duarte 1999 (2005). Two phylogenetically significant new Calycopis species (Lycaenidae: Theclinae: Eumaeini). Journal of Research on the Lepidoptera 38:27-34.

Romdal, T. S., R. K. Colwell, and C. Rahbek. 2005. The influence of band sum area, domain extent and range sizes on the latitudinal mid-domain effect. Ecology 86:235-244.

Rota, J. 2005. Larval and pupal descriptions of the Neotropical choreutid genera Rhobonda Walker and Zodia Heppner (Lepidoptera: Choreutidae). Annals of the Entomolical Society of America 98: 37-47.

Sharkey, M. J. and R. A. Briceño G. 2005. Review of Sesioctonus Viereck (Hymenoptera: Braconidae: Agathidinae) from Colombia with descriptions of five new species. Zootaxa 1002:45-58.

Wagner, D. L. 2005. Caterpillars of Eastern North America: A Guide to Identification and Natural History. Princeton University Press. 512 pp.


Brown, B. V. 2004. Revision of the subgenus Udamochiras of Melaloncha bee-killing flies (Diptera: Phoridae: Metopininae). Zoological Journal of the Linnean Society. 140: 1-42.

Colwell, R. K. 2004. Biota 2: The Biodiversity Database Manager. Sinauer Associates, Sunderland, MA. Software application and 860 page manual.

Colwell, R. K., C. X. Mao, and J. Chang. 2004. Interpolating, extrapolating, and comparing incidence-based species accumulation curves. Ecology 85:2717-2727.

Colwell, R. K., C. Rahbek, N. J. Gotelli. 2004. The mid-domain effect and species richness patterns: What have we learned so far? American Naturalist 163:E1-E23.

Dubovikoff, D. A., and J. T. Longino. 2004. A new species of the genus Bothriomyrmex Emery, 1869 (Hymenoptera: Formicidae: Dolichoderinae) from Costa Rica. Zootaxa 776:1-10.

Erwin, T. L. 2004. The beetle family Carabidae of Costa Rica and Panamá: Descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa 537: 1-18.

Gillespie, J. J., Kjer, K. M., Riley, E. G., and Tallamy, D. W. 2004. The Evolution of Cucurbitacin Pharmacophagy in Rootworms: insight from Luperini Paraphyly. Pp 37-57, in New Developments In The Biology Of Chrysomelidae. Edited by Pierre H. Jolivet (Paris ), Jorge A. Santiago-Blay (Washington) and Michael Schmitt (Bonn).

Hamilton, R.W. and A.L. Novinger. 2004. Eugnamptine weevils of La Selva, Costa Rica (Coleoptera: Rhynchitidae). Zootaxa 610:1-43.

Hansson, C. 2004. Eulophidae of Costa Rica, 2. Memoirs of the American Entomological Institute 75:1-537.

Jetz, W., C. Rahbek, and R. K. Colwell. 2004. The coincidence of rarity and richness and the potential signature of history in centres of endemism. Ecology Letters 7:1180-1191.

Klimov, P. B., M. Lekveishvili, A. P. G. Dowling, and B. M. OConnor. 2004. Multivariate analysis of morphological variation in two cryptic species of Sancassania (Acari: Acaridae) from Costa Rica. Annals of the Entomological Society of America 97:322-345.

Koh, L. P., R. R. Dunn, N. S. Sodhi, R. K. Colwell, H. C. Proctor, and V. S. Smith. 2004. Species coextinctions and the biodiversity crisis. Science 305:1632-1634.

LaPolla, J.S. 2004. Acropyga (Hymenoptera: Formicidae) of the World. Contributions to the American Entomological Institute 33:1-130.

Longino, J. T., and S. Cover. 2004. A Revision of the Aphaenogaster phalangium complex (Hymenoptera: Formicidae: Myrmicinae). Zootaxa 655:1-12.

Martins, U. R., and M. H. M. Galileo. 2004. Contribuição ao conhecimento dos Hemilophini (Coleoptera, Cerambycidae, Lamiinae), principalmente da Costa Rica. Revista Brasileira de Entomologia 48:467-472.

Noyes, J.S. 2004. Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae). Memoirs of the American Entomological Institute 73:1-459.

Saporito, R. A., H. M. Garraffo, M. A. Donnelly, A. L. Edwards, J. T. Longino, and J. W. Daly. 2004. Formicine ants: An arthropod source for the pumiliotoxin alkaloids of dendrobatid poison frogs. Proceedings of the National Academy of Sciences, USA 101:8045-8050.

Schonberg, L., J. T. Longino, N. M. Nadkarni, S. P. Yanoviak, and J. C. Gering. 2004. Arboreal Ant Species Richness in primary, secondary forest and pasture habitats of a tropical montane landscape. Biotropica 36:402-409.


Anderson, R.S. 2003. Neotropical Dryophthoridae: Redescription of the genus Melchus Lacordaire with description of Daisya Anderson, new genus, and seven new species (Coleoptera; Curculionoidea). The Coleopterists Bulletin 57: 413-432.

Behan-Pelletier, V. M., B. Eamer 2003. Redefinition of Pelopsis (Acari: Oribatida: Mycobatidae), with description of Pelopsis baloghi sp. n. from Costa Rica. Acta Zoologica Academiae Scientiarum Hungaricae 49:5-15.

Briceño, R. A. 2003. Taxonomic revision of the genus Sesioctonus Viereck (Hymenoptera: Braconidae: Agathidinae). Journal of Hymenoptera Research 12:238-271.

Brown, J. W. & K. Nishida. 2003. First record of larval endophagy in Euliini (Tortricidae: Euliini): A new species of Seticosta from Costa Rica. Journal of the Lepidopterists' Society 56:113-120.

Brown, J. W. 2003. An illustrated guide to the Orthocomotis Dognin (Tortricidae) of Costa Rica, with summaries of their spatial and temporal distribution. Journal of the Lepidopterists' Society 57:253-269.

Davis, D.R. 2003. A Monograph of the Family Arrhenophanidae (Lepidoptera: Tineoidea). Smithsonian Contributions to Zoology 620:1-80, 255 figs., 9 maps.

Fernández C., F. 2003. Revision of the myrmicine ants of the Adelomyrmex genus-group (Hymenoptera: Formicidae). Zootaxa 361:1-52.

Fernández C., F., and W. P. MacKay. 2003. The Myrmicine ants of the Adelomyrmex laevigatus species complex (Hymenoptera: Formicidae). Sociobiology 41:1-12.

Furth, D. G., J. T. Longino, and M. Paniagua. 2003. Survey and quantitative assessment of flea beetle diversity in a Costa Rican rainforest (Coleoptera: Chrysomelidae: Alticinae). Pp. 1-23 in D. G. Furth, editor. Topics in Leaf Beetle Biology: Proceedings of the Fifth International Symposium on the Chrysomelidae, 25-27 August 2000, Iguassu Falls, Brazil. XXI International Congress of Entomology. Pensoft Publishers, Moscow.

Hansson, C., and J. LaSalle 2003. Revision of the Neotropical species of the tribe Euderomphalini (Hymenoptera: Eulophidae). Journal of Natural History 37:697-778.

Kaczmarek, L. 2003. New records of and key to Tardigrada from Costa Rica. Zootaxa 177:1-4.

Leathers, J.W. and M.J. Sharkey 2003. Taxonomy and life history of Costa Rican Alabagrus (Hymenoptera: Braconidae), with a key to world species. Contributions in Science 497:1-78.

Longino, J. T. 2003 The Crematogaster (Hymenoptera, Formicidae, Myrmicinae) of Costa Rica. Zootaxa 151:1-150.

Longino, J. T. 2003. A new Costa Rican species of Leptanilloides (Hymenoptera: Formicidae: doryline section: Leptanilloidinae). Zootaxa 264:1-6.

Niedbala, W. 2003. Ptyctimous mites (Acari: Oribatida) of Costa Rica. Annales Zoologici 53:259-334.

Prena J. 2003. The Middle American species of Ambates Schönherr (Coleoptera, Curculionidae, Baridinae). Beiträge zur Entomologie 53:167-198.

Prena J. 2003. The Pardisomus species from Costa Rica, with descriptions of four new species and one larva (Coleoptera, Curculionidae, Baridinae). Beiträge zur Entomologie 53:199-216.

Smith, D. R. and D. H. Janzen. 2003. Food plants and life histories of sawflies of the families Tenthredinidae and Pergidae (Hymenoptera) in Costa Rica, with descriptions of four new species. Journal of Hymenoptera Research 12: 312-332.

Wilson, E. O. 2003. Pheidole in the New World A Dominant, Hyperdiverse Ant Genus. Harvard University Press, Cambridge Mass. 818 pgs.


Anderson, R. S. 2002. The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus , Mesocordylus , Metamasius and Rhodobaenus (Coleoptera; Curculionoidea). Zootaxa 80: 1-94.

Baker, A. S. and Lindquist, E. E. 2002. Aethosolenia laselvensis gen. nov., sp. nov., a new eupodoid mite from Costa Rica (Acari: Prostigmata). Systematic and Applied Acarology Special Publications 11:1-11.

Carpenter, J. M. and J. Kojima. 2002. A new species of paper wasp from Costa Rica (Hymenoptera: Vespidae, Polistinae, Epiponini). Journal of the New York Entomological Society 110: 212-223.

Gauld, I., Sithole, R., Ugalde Gómez, J., Godoy, C. 2002. Ichneumonidae of Costa Rica, 4. Memoirs of the American Entomological Institute 66. 768pp.

Goldarazena, A., R. Ochoa, C. C. Childers 2002. Systematic revision of the genus Deleonia (Acari: Tarsonemidae). International Journal of Acarology 28:221-240.

Hansson, C. 2002. Eulophidae of Costa Rica, 1. Memoirs of the American Entomological Institute 67:1-290. Hespenheide, H. A. 2002. A review of North and Central American Paragrilus Saunders, 1871 (Coleoptera: Buprestidae: Agrilinae). Zootaxa 43:1-28.

Hespenheide, H. A. 2002. A review of North and Central American Paragrilus Saunders, 1871 (Coleoptera: Buprestidae: Agrilinae). Zootaxa 43:1-28.

Hespenheide, H.A. and L.M. LaPierre. 2002. A review of Ptous (Coleoptera: Curculionidae: Cryptorhynchinae). Proceedings of the Entomological Society of Washington.104:856-862.

Longino, J., R. K. Colwell, and J. A. Coddington 2002. The ant fauna of a tropical rainforest: estimating species richness three different ways. Ecology 83:689-702.

Longino, J. T., and R. R. Snelling. 2002. A taxonomic revision of the Procryptocerus (Hymenoptera: Formicidae) of Central America. Contributions in Science (Los Angeles County Museum of Natural History) 495:1-30.

Marsh, P. M. 2002. The Doryctinae of Costa Rica (excluding the genus Heterospilus). Memoirs of the American Entomological Institute 70:1-319.

Rifkind, J. 2002. New Central American and Mexican Enoclerus (Coleoptera: Cleridae). Natural History Museum of Los Angeles County, Contributions in Science 487:1-16.

Solís, A. y B. Kohlmann, B. 2002. El género Canthon (Coleoptera: Scarabaeidae) en Costa Rica. Giornale italiano di entomologia 10:1-68.


Andrade, M. L. (de) 2001. A remarkable Dominican amber species of Cylindromyrmex with Brazilian affinities and additions to the generic revision (Hymenoptera: Formicidae). Beiträge zur Entomologie 51:51-63

Brown, B. V. 2001. Taxonomic revision of Neodohrniphora, subgenus Eibesfeldtphora (Diptera: Phoridae). Insect Systematics and Evolution 32:393-409.

Brown, B. V. 2001. Biodiversity of flies, gnats and mosquitoes. In S. A. Levin, ed. Encyclopedia of biodiversity. Academic Press, NY.

Coddington, J. A., and R. K. Colwell. 2001. Arachnids, Pages 199-218 in S.Levin, ed. Encyclopedia of biodiversity. NY, Academic Press.

Colwell, R., J. Longino, D. Brenes, N. Oconitrillo, M. Paniagua, R. Vargas. 2001. Los artrópodos: protagonistas en la estimación y conservación de la biodiversidad. Recuadro 3.4 en R. Primack, R. Rozzi, P. Feinsinger, R. Dirzo, F. Massardo. Conservación biologica. Fondo de Cultura Económica, México.

Gilbert, A. J., F. E. Andrews, & C. L. Staines. 2001. An unusual host record for Imatidium and new distributional records for I. rufiventre Boh. and I. thoracicum Fab. (Coleoptera: Chrysomelidae: Cassidinae). Pan-Pacific Entomologist 77:278-280.

Goldarazena, A., R. Ochoa, R. Jordana and B. M. OConnor 2001. Revision of the genus Adactylidium (Acari: Acarophenacidae). Proceedings of the Entomological Society of Washington 103:473-516.

Gotelli, N. J., and R. K. Colwell 2001. Quantifying biodiversity: Procedures and pitfalls in the measurement and comparison of species richness. Ecology Letters 4:379-391.

Hespenheide, H. A. 2001. Beetles. Pages 351-358, in S. A. Levin, ed. Encyclopedia of biodiversity. Academic Press, NY.

Kohlmann, B. y A. Solís. 2001. El género Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale italiano di entomologia. 9:159-261.

Landry, B., and J. A. Powell. 2001. Systematics and Phylogeny of Sparganothina and Related Taxa (Lepidoptera: Tortricidae: Sparganothini). UC Publications in Entomology, 121. University of California Press, Berkeley, CA, USA.

Levin, S., (Ed.) R. K. Colwell, G. Daily, L. J., H. A. Mooney, E. Schulze, and D. Tilman. (Assoc. Eds.) 2001. Encyclopedia of biodiversity, Pages 4666. NY, Academic Press.

Lindquist, E. E. 2001. Poising for a new century: diversification in acarology. Pp. 17-34 in R.B. Halliday, et al. (Editors), Acarology: Proceedings of the 10th International Congress. 1998, Canberra, Australia. CSIRO Publishing, Melbourne.

Morehead, S., J. Seger, D.H. Feener, Jr. and B.V. Brown. 2001. Evidence for a cryptic species complex in the ant parasitoid Apocephalus paraponerae (Diptera: Phoridae). Evolutionary Ecology Research. 3: 273-284.

Prena, J. 2001. A revision of the neotropical weevil genus Pantoteles Schönherr (Coleoptera, Curculionidae, Baridinae). Trans. Am. Entomol. Soc. 127:305-358.

Wagner, D. L. 2001. Las granjas de marisposas: un colorido ejemplo de manejo sustentable. In. Elementos de Conservacion Biologica: Perspectivas Latinoamericanas. R Primack, R. Rozzi, P. Feinsinger, R. Dirzo, and F. Massardo (eds.) Fondo de Cultura Economica, Ciudad de Mexico, Mexico.

Wagner, D. L. 2000. The Biodiversity of Moths, pp. 249-270. In the Encyclopedia of Biodiversity, S. Levin et al. (eds.) Academic Press. San Diego, CA.


Brown, B.V. 2000. Revision of the "Apocephalus miricauda-group" of ant-parasitizing flies (Diptera: Phoridae). Contributions in Science. 482:1-62.

Colwell, R. K. 2000. A barrier runs through it...or maybe just a river (Commentary). PNAS 97:13470-13472.

Colwell, R. K. 2000. Hummingbirds and flower mites: Contrasts and conflicts between two nectarivores, Pages 65-91 in M. A. S. Alves, J. M. C. Silva, M. Van Sluys, H. G. Bergallo, and C. F. D. Rocha, eds. A ornitologia no Brasil: Pesquisa atual e perspectivas. Anais do VII Congresso Brasileiro de Ornitologia. Río de Janeiro, ED UERJ, Universidade do Estado do Rio de Janeiro.

Colwell, R. K. 2000. Rensch's Rule crosses the line: Convergent allometry of sexual size dimorphism in hummingbirds and flower mites. American Naturalist 156:495-510.

Colwell, R. K., and D. C. Lees. 2000. The mid-domain effect: geometric constraints on the geography of species richness. Trends in Ecology and Evolution 15:70-76.

Colwell, R. K., and D. C. Lees. 2000. Reply from R. K. Colwell and D. C. Lees. Trends in Ecology and Evolution 15:289.

Corrales, J. F., and M. E. Epstein. 2000. New species in the New World Natada complex (Lepidoptera: Limacodidae). Revista de Biologia Tropical 48:591-604.

Gauld, I. (2000) Ichneumonidae of Costa Rica III: Introduction and keys to the species of the subfamilies Brachycrtinae, Cremastinae, Labeninae, Oxytorinae. Memoirs of the American Entomological Institute 63. 410 pp.

Johnson, P. J. 2000. New species and records of Platycrepidius Candeze from Costa Rica and Panama (Coleoptera: Elateridae). Coleop. Bull. 54:284-290.

LaPierre, L. M. 2000. Prey selection and diurnal activity of Holcocephala oculata (F.) (Diptera: Asilidae) in Costa Rica. Proceedings of the Entomological Society of Washington 102: 643-651.

Longino, J. T. 2000. The ants of Monteverde. In N. M. Nadkarni and N. T. Wheelwright (Ed.). Monteverde: ecology and conservation of a tropical cloud forest, pp. 132-136. Oxford University Press, New York, New York.

Longino, J. T. 2000. Myrmecophytes. In N. M. Nadkarni and N. T. Wheelwright (Ed.). Monteverde: ecology and conservation of a tropical cloud forest, pp. 291-293. Oxford University Press, New York, New York.

Longino, J. T. 2000. What to do with the data. In D. Agosti, J. D. Majer, L. E. Alonso and T. R. Schultz (Ed.). Measuring and monitoring biological diversity: standard methods for ground-living ants, pp. 186-203. Smithsonian Institution Press, Washington, D.C., USA.

Naskrecki, P. 2000. Katydids of Costa Rica. Volume I. Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato). The Orthopterists' Society at the Academy of Natural Sciences of Philadelphia. Publications on Orthopteran Diversity. iv+164pp, CD-ROM.

Ochoa, R. and B. M. OConnor. 2000 Revision of the genus Horstiella (Acari: Acaridae), mites associated with Neotropical Epicharis bees (Hymenoptera: Apidae). Annals of the Entomological Society of America. 93: 713-737.

Oliver, I., A. Pik, D. Britton, J. M. Dangerfield, R. K. Colwell, and A. J. Beattie. 2000. Virtual biodiversity assessment systems: the application of bioinformatics technologies to the accelerated accumulation of biodiversity information. BioScience 50:441-450.

Staines, C. L. 2000. A new species of Xenochalepus Weise, 1910 (Coleoptera: Chrysomelidea: Hispinae) from Mesoamerica. Journal of the New York Entomological Society 108:95-97.


Brown, B.V. 1999. Differential host use by neotropical phorid flies (Diptera: Phoridae) that are parasitoids of ants (Hymenoptera: Formicidae). Sociobiology. 33: 95-103.

Chazdon, R. L., R. K. Colwell, and J. S. Denslow. 1999. Tropical tree richness and resource-based niches. Science 285:1459.

Colwell, R. K. 1999. RangeModel: a Monte Carlo simulation tool for assessing geometric constraints on species richness. (Software and User's Guide.) Published at

Colwell, R. K., and S. Naeem. 1999. Sexual sorting in hummingbird flower mites. Annals of Entomological Society of America 92: 952-959.

Colwell, R. K., and P. Naskrecki. 1999. The evolution of host-plant affiliation, body size and sexual dimorphism in hummingbird flower mites (Ascidae: Rhinoseius and Tropicoseius). Pp. 79-86 in G. R. Needham, R. Mitchell, D. J. Horn and W. C. Welbourn, eds. Acarology IX, Volume 2: Symposia (IXth International Congress of Acarology ). Ohio Biological Survey, Columbus, Ohio.

Denmark, H., G. A. Evans, H. Aguilar, C.Vargas, and R. Ochoa. 1999. Phytoseiidae of Central America (Acari: Mesostigamta). Indira Publishing House, Michigan, USA. 125 pp.

Hespenheide, H. A., and V. Dang. 1999. Biology and ecology of leaf-mining Hispinae (Coleoptera, Chrysomelidae) of the La Selva Biological Station, Costa Rica. Pp. 375-389 In M. L. Cox (Ed.). Advances in Chrysomelidae Biology 1, pp. Backhuys Publishers, Leiden, The Netherlands.

Naskrecki, P. and D. Otte. 1999. An illustrated catalog of Orthoptera. Vol. I. Tettigonioidea. The Orthopterists' Society, Academy of Natural Sciences of Philadelphia. (CD ROM. Approximately 3000 pages if printed.)

Staines, C. 1999. Possible mimetic complexes in Central American Cephaloleia (Coleoptera: Chrysomelida: Hispinae). Pp. In M. L. Cox (Ed.). Advances in Chrysomelidae Biology 1, pp. 239-246. Backhuys Publishers, Leiden, The Netherlands.

Staines, C. L. 1999(1998). A review of the genus Physocoryna Guerin-Meneville, 1844 (Coleoptera: Chrysomelidae: Hispinae). J. New York Ent. Soc. 106:163-169.


Behan-Pelletier, V. M. 1998. Ceratozetoidea (Acari: Oribatida) of lowland tropical rainforest, La Selva, Costa Rica. Acarologia 39: 349-381.

Brown, B. V., and D. H. Feener Jr. 1998. Parasitic phorid flies (Diptera: Phoridae) associated with army ants (Hymenoptera: Formicidae: Ecitoninae, Dorylinae) and their conservation biology. Biotropica 30:482-487.

Chazdon, R. L., R. K. Colwell, J. S. Denslow, and M. R. Guariguata. 1998. Statistical methods for estimating species richness of woody regeneration in primary and secondary rain forests of northeastern Costa Rica. Pp. In F. Dallmeier and J. A. Comiskey (Ed.). Forest Biodiversity Research, Monitoring and Modeling: Conceptual Background and Old World Case Studies, pp. 285-309. Parthenon Publishing, Paris, France.

Halliday, R. B., D. E. Walter, and E. E. Lindquist. 1998. Revision of the Australian Ascidae (Acari: Mesostigmata). Invertebrate Taxonomy 12:1-54.

Heraty, J. M. 1998. The genus Dilocantha (Hymenoptera: Eucharitidae). Proc. Entomol. Soc. Washington 100:72-87.

Jordal, B. H. 1998. New species of Scolytodes (Coleoptera: Scolytidae) from Costa Rica and Panamá. Revista de biología tropical 46:407-420. Kissinger, D. G. 1998. Apioinidae from North and Central America. Part 5. Description of genus Apionion and 4 new species (Coleoptera). Insecta Mundi 12:93-99.

Kissinger, D. G. 1998. Apioinidae from North and Central America. Part 5. Description of genus Apionion and 4 new species (Coleoptera). Insecta Mundi12:93-99.

Naskrecki, P., and R. K. Colwell. 1998. Systematics and host plant affiliations of hummingbird flower mites of the genera Tropicoseius Baker and Yunker and Rhinoseius Baker and Yunker (Acari: Mesostigmata: Ascidae). Thomas Say Publications in Entomology: Monographs. Entomological Society of America. 185 pp.

Ochoa, R., and B. M. Oconnor. 1998. Two new species of the genus Excelsotarsonemus (Acari: Tarsonemidae). Internat. J. Acarol. 24(3):179-187.

Staines, C. L. 1998. A new species of Cephaloleia Chevrolat (Coleoptera: Chrysomelidae: Hispinae) from Costa Rica. Proc. Ent. Soc. Wash. 100:672-673.

Thunes, K. H. 1998. Bark and ambrosia beetles (Coleoptera: Curculionidae, Scolytinae and Platypodinae) in a neotropical rainforest. Comparing occurrence and distribution between different forest habitats within a continuous reserve in Costa Rica. Institute of Zoology, University of Bergen, Norway.


Behan-Pelletier, V. M. 1997. The semi-aquatic genus Tegeocranellus (Acari: Oribatida: Ameronothroidea) of North and Central America. Canadian Entomologist 129:537-577.

Brown, B. V. 1997. Parasitic phorid flies: a previously unrecognized cost to aggregation behavior of male stingless bees. Biotropica 29:370-372.

Brown, B. V. 1997. Revision of the Apocephalus attophilus-group of ant-decapitating flies (Diptera: Phoridae). Contributions in Science 468:1-60.

Gauld, I. 1997. Ichneumonidae of Costa Rica II: Introduction and keys to the species of the subfamilies Anomaloninae, Ctenopelmatinae, Diplazontinae, Lycorininae, Phrudinae, Tryphoninae, Xoridinae. Memoirs of the American Entomological Institute 57. 585 pp.

Kohlmann, B. y A. Solís. 1997. El género Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale italiano di entomologia. 8:343-382. Longino, J. T., and R. K. Colwell. 1997. Biodiversity assessment using structured inventory: capturing the ant fauna of a lowland tropical rainforest. Ecological Applications 7:1263-1277.

Naskrecki, P. 1997. A revision of the neotropical genus Acantheremus Karny, 1907 (Orthoptera: Tettigoniidae: Copiphorinae). Transactions of the American Entomological Society 123:137-161.

Nickle, D. A. and P. Naskrecki 1997. Recent developments in the systematics of Tettigoniidae and Gryllidae. In: The bionomics of grasshoppers, katydids and their kin, S. K. Gangwere, M. C. Muralirangan and Meera Muralirangan, Ed., CAB International, pp. 41-58.

Ochoa, R, C.Vargas, D. Walter,.and B. M. OConnor. 1997. Two new species of the genus Ceratotarsonemus (Acari: Tarsonemidae). Internat. J. Acarol. 23(3): 177-183.

OConnor, B. M., R. K. Colwell, and S. Naeem. 1997. The flower mites of Trinidad III. The genus Rhinoseius (Acari: Ascidae). Miscellaneous Publication No. 184. University of Michigan Museum of Zoology.

Sharkov, A., and J. B. Woolley. 1997. A revision of the genus Hambletonia Compere (Hymenoptera: Encyrtidae). Journal of Hymenoptera Research 6:191-218.

Zitani, N. M., S. R. Shaw, and D. H. Janzen. 1997. Description and biology of a new species of Meteorus Haliday (Hymenoptera: Braconidae, Meteorinae) from Costa Rica, parasitizing larvae of Papilio and Parides (Lepidoptera: Papilionidae). Journal of Hymenoptera Research 6:178-185.


Ayala, F. J., J. K. Wetterer, J. T. Longino, and D. L. Hartl. 1996. Molecular phylogeny of Azteca ants (Hymenoptera: Formicidae) and the colonization of Cecropia trees. Molecular Phylogenetics and Evolution 5:423-428.

Brown, B. V. 1996. Preliminary analysis of a host shift: revision of the Neotropical species of Apocephalus, subgenus Mesophora (Diptera: Phoridae). Contributions in Science 462:1-36.

Colwell, R. K. 1996-1999. Biota: The biodiversity database manager. Sinauer Associates, Sunderland, MA. [Manual, 1996, 574 pp.; Manual Supplement 1, 1997, 66 pp.; Manual Supplement 2, 1998, 18 pp.; Software copyright 1996, 1997, 1998, 1999.]

Hespenheide, H. A. 1996. Chrysomelidae of the subfamily Clytrinae as models for mimicry complexes. Pp. 227-239 In P. H. A. Jolivet and M. L. Cox (Ed.). Chrysomelidae Biology, Vol. 2, Ecological Studies. SPB Academic Publishing, Amsterdam, The Netherlands.

Hespenheide, H. A. 1996. The role of plants in structuring communities of mimetic insects. Pp. 109-126 In A. C. Gibson (Ed.). Neotropical Diversity and Conservation. Mildred Mathias Botanical Garden, University of California, Los Angeles, Los Angeles, CA, USA.

Longino, J. T. 1996. Taxonomic characterization of some live-stem inhabiting Azteca (Hymenoptera: Formicidae) in Costa Rica, with special reference to the ants of Cordia (Boraginaceae) and Triplaris (Polygonaceae). Journal of Hymenoptera Research 5:131-156.

Shaw, S. R. 1996. Plynops, a peculiar new neotropical genus and seven new species (Hymenoptera: Braconidae: Euphorinae). Journal of Hymenoptera Research 5:166-183.

Staines, C. L. 1996. The genus Cephaloleia (Coleoptera: Chrysomelidae) in Central America and the west Indies. Special Publication No. 3 of the Revista de Biologia Tropical. 87 pp.


Brown, B. V., and D. H. Feener Jr. 1995. Efficiency of two mass sampling methods for sampling phorid flies (Diptera: Phoridae) in a tropical biodiversity survey. Contributions in Science 459:1-10.

Colwell, R. K. 1995. Effects of nectar consumption by the hummingbird flower mite Proctolaelaps kirmsei on nectar availability in Hamelia patens. Biotropica 27:206-217.

Colwell, R. K., and J. A. Coddington. 1995. Estimating terrestrial biodiversity through extrapolation. Pp. 101-118 in D. L. Hawksworth, eds. Biodiversity: Measurement and estimation. Chapman and Hall, London.

Husband, R.W., R. Ochoa, and P. Naskrecki. 1995. Buprestapolipus megachelus n. gn., n. sp. (Acari: Podapolipidae) from Euchroma gigantea L. (Coleoptera: Buprestidae). Internat. J. Acarol. 21(3):185-194.

Infante, F., P. Hanson, and R. Wharton. 1995. Phytophagy in the genus Monitoriella (Hymenoptera: Braconidae) with description of new species. Ann. Entomol. Soc. Am. 88:406-415.

Longino, J. T., and P. Hanson. 1995. The ants (Formicidae). Pp. In P. Hanson and I. Gauld (Ed.). The Hymenoptera of Costa Rica, pp. 588-620. Oxford University Press, Oxford, U.K.

Longino, J. T., and D. A. Hartley. 1995 (1994). Perissomyrmex snyderi (Hymenoptera, Formicidae) is native to Central America and exhibits worker polymorphism. Psyche 101:195-202.

Naskrecki, P., and R. K. Colwell. 1995. A new genus and two new species of Melicharini from Venezuela (Acari: Mesostigmata: Ascidae). Annals of the Entomological Society of America 88:284-293.

Ochoa, R., P. Naskrecki, and R. K. Colwell. 1995. Excelsotarsonemus kaliszewskii, a new genus and new species from Costa Rica (Acari: Tarsonemidae). International Journal of Acarology 21:67-74.


Longino, J. T. 1994. How to measure arthropod diversity in a tropical rainforest. Biology International 28:3-13.

Colwell, R. K. 1994-1999. EstimateS: statistical estimation of species richness and shared species from samples (Software and User's Guide). Published at

Colwell, R. K., and J. A. Coddington. 1994. Estimating terrestrial biodiversity through extrapolation. Phil. Trans. R. Soc. Lond. B 345:101-118.

Colwell, R. K., and G. C. Hurtt. 1994. Nonbiological gradients in species richness and a spurious Rapoport effect. American Naturalist 144:570-595.

Hespenheide, H. A. 1994. An overview of faunal studies. In L. McDade, K. S. Bawa, G. S. Hartshorn and H. A. Hespenheide (eds.), La Selva: Ecology and History of a Neotropical Rainforest, pp. University of Chicago Press, Chicago, IL, USA.

McDade, L. A., K. S. Bawa, H. A. Hespenheide, and G. S. Hartshorn, ed. 1994. La Selva, Ecology and Natural History of a Neotropical Rainforest. University of Chicago Press, Chicago, IL, USA.

Newstrom, L. E., G. W. Frankie, H. G. Baker, and R. K. Colwell. 1994. Diversity of long-term flowering patterns at La Selva. In L. A. McDade, K. S. Bawa, G. S. Hartshorn and H. A. Hespenheide, eds. La Selva: ecology and natural history of a lowland tropical rain forest. University of Chicago Press, Chicago.


Colwell, R. K. 1993. Management of wildland biodiversity as a resource for biotechnology. Pp. 130-134 in W. Tien, S.-F. Chen, L. Lo and Y. T. Shyu, eds. Biotechnology: Theory and applications 1993. Development Center for Biotechnology, Tapei, R. O. C.

Colwell, R. K., and S. Naeem. 1993. Life history patterns of hummingbird flower mites in relation to host phenology and morphology. Pp. 23-44 in M. A. Houck, eds. Mites: Ecological and evolutionary analyses of life history patterns. Chapman and Hall, New York.

Kremen, C., R. K. Colwell, T. L. Erwin, D. D. Murphy, R. F. Noss, and M. A. Sanjayan. 1993. Terrestrial arthropod assemblages: their use in conservation planning. Conservation Biology 7:796-808.

Lindquist, E. E., and M. L. Moraza. 1993. Pyrosejidae, a new family of trigynaspid mites (Acari: Mesostigmata: Cercomegistina) from Middle America. Acarologia 34:283-307.

Longino, J. T. 1993. The art and science of taxonomy. National Geographic Research 9(1):80-85.

Marsh, P. M. 1993. Description of new Western Hemisphere genera of the subfamily Doryctinae (Hymenoptera: Braconidae). Contributions of the American Entomological Institute 28:1-58.


Colwell, R. K. 1992. Human aspects of biodiversity: An evolutionary perspective. International Union of Biological Sciences, Monograph No. 8. Pp. 209-222 in O. T. Solbrig, H. M. van Emden and P. G. W. J. van Oordt, eds. Biological diversity and global change. IUBS Press, Paris

Colwell, R. K. 1992. Making sense of ecological complexity: A personal and conceptual retrospective. Biotropica 24(2b):226-232.

Hespenheide, H. A. 1992. A review of the genus Tachygonus Schoenherr (Coleoptera: Curculionidae) in America north of Mexico. Proc. Ent. Soc. Wash. 94:1-11.

Hespenheide, H. A., and C. Kim. 1992. Clutch size, survivorship and larval biology of Pachyschelus psychotriae Fisher (Coleoptera: Buprestidae). Ann. Entomol. Soc. Amer. 85:48-52.